← Back to course dashboard 🔬 Module V · Analytical & Biochemical Techniques
UNIT 5.2

Spectrophotometry & Biomolecule Quantification

Measuring light absorption to determine concentration

🎯 After this unit, you will be able to:

  • Explain the principle of spectrophotometry
  • Apply the Beer-Lambert law to calculate concentrations
  • Perform common colorimetric assays for proteins, sugars, and pigments
  • Construct and use a standard curve

📊 What is Spectrophotometry?

Spectrophotometry is a technique that measures how much light a substance absorbs at different wavelengths. Since many biomolecules absorb light in predictable ways, spectrophotometry is one of the most widely used tools in biochemistry labs .

Key insight: The amount of light absorbed is directly proportional to the concentration of the absorbing molecule. This allows us to quantify unknown concentrations by comparing to known standards .

🔬 [Diagram: Basic spectrophotometer components: light source, monochromator, sample cuvette, detector — to be inserted]

📐 The Beer-Lambert Law

The relationship between light absorption and concentration is described by the Beer-Lambert law:

A = ε × c × l

Where:

  • A = Absorbance (no units)
  • ε = Molar extinction coefficient (M⁻¹ cm⁻¹) — a constant for each molecule at a specific wavelength
  • c = Concentration (M or other units)
  • l = Path length (cm) — usually 1 cm for standard cuvettes

Since path length is fixed (1 cm), and ε is known for many compounds, we can calculate concentration directly from absorbance:

c = A / (ε × l)

Important Notes

  • Absorbance is directly proportional to concentration — a linear relationship
  • This relationship holds only at low to moderate concentrations
  • At high concentrations, deviations occur (too much light is absorbed)
  • Always work within the linear range of the assay
🔍 Did you know? The Beer-Lambert law is named after August Beer (German mathematician) and Johann Heinrich Lambert (Swiss mathematician), who described the relationship between light absorption and concentration in the 18th and 19th centuries .

📈 Absorbance vs. Transmittance

Spectrophotometers measure the intensity of light before (I₀) and after (I) passing through a sample. Two related terms are used:

  • Transmittance (T): T = I / I₀. Usually expressed as %T = (I/I₀) × 100
  • Absorbance (A): A = -log₁₀(T) = log₁₀(I₀/I)

Absorbance is preferred because it's linearly related to concentration. Transmittance is exponential .

📊 [Graph: Relationship between transmittance and absorbance — to be inserted]

📉 Standard Curves: The Key to Quantification

When ε is unknown, or when using colorimetric assays where the color intensity depends on reaction conditions, we use a standard curve.

How to Construct a Standard Curve

  1. Prepare a series of standards with known concentrations
  2. Measure absorbance of each standard
  3. Plot absorbance vs. concentration
  4. Add a line of best fit (linear regression)
  5. The equation of the line (y = mx + b) allows you to calculate unknown concentrations from absorbance
📈 [Diagram: Typical standard curve with linear regression — to be inserted]

✅ Tips for Good Standard Curves

  • Use at least 5-6 standard concentrations
  • Cover the expected range of your unknowns
  • Include a blank (zero concentration)
  • R² should be > 0.98 for a good fit
  • Never extrapolate beyond your highest standard

🧪 Bradford Protein Assay Standard Curve

To quantify unknown protein samples, you prepare BSA standards at 0, 0.1, 0.2, 0.4, 0.6, 0.8, 1.0 mg/mL. After adding Bradford reagent and measuring A₅₉₅, you get a standard curve with equation y = 0.85x + 0.02 (R² = 0.99). An unknown sample gives A₅₉₅ = 0.45. What is its protein concentration?

Solution: 0.45 = 0.85x + 0.02 → x = (0.45 - 0.02)/0.85 = 0.51 mg/mL

🧪 Common Colorimetric Assays in Plant Biochemistry

🥚

Bradford Protein Assay

595 nm (blue)

Coomassie Brilliant Blue G-250 binds to proteins, shifting absorbance from 465 nm to 595 nm. Fast, sensitive, compatible with many reagents .

🍬

Phenol-Sulfuric Acid (Total Sugars)

490 nm (yellow-orange)

Concentrated H₂SO₄ hydrolyzes sugars to furfural derivatives, which react with phenol to form colored compounds. Measures total carbohydrates .

🍇

Folin-Ciocalteu (Total Phenolics)

765 nm (blue)

Phenols reduce phosphomolybdic/phosphotungstic acid in alkaline conditions, producing a blue color. Standardized with gallic acid .

🌿

Chlorophyll Extraction

664 nm, 647 nm

Chlorophyll a and b absorb at specific wavelengths in 80% acetone. Equations allow calculation of concentrations .

🍋

Vitamin C (Ascorbic Acid)

520 nm

2,6-dichlorophenolindophenol (DCPIP) reduction; color change from blue to colorless .

🔵

DNSA Reducing Sugars

540 nm

3,5-dinitrosalicylic acid reacts with reducing sugars to form colored compounds .

📋 Detailed Assay Protocols

Bradford Protein Assay Protocol

Step Procedure
1 Prepare BSA standards: 0, 0.1, 0.2, 0.4, 0.6, 0.8, 1.0 mg/mL
2 Add 100 μL of each standard or unknown to labeled test tubes
3 Add 5 mL Bradford reagent to each tube, vortex
4 Incubate at room temperature for 5-10 minutes
5 Measure absorbance at 595 nm
6 Construct standard curve and calculate unknown concentrations

Chlorophyll Extraction and Quantification

Arnon's equations (in 80% acetone):

Chlorophyll a (mg/L) = 12.7 × A₆₆₃ - 2.69 × A₆₄₇
Chlorophyll b (mg/L) = 22.9 × A₆₄₇ - 4.68 × A₆₆₃
Total chlorophyll = 20.2 × A₆₄₇ + 8.02 × A₆₆₃

Procedure:

  1. Extract leaf tissue (0.1 g) in 10 mL 80% acetone
  2. Centrifuge or filter to remove debris
  3. Measure absorbance at 647 and 663 nm
  4. Apply equations to calculate concentration
🍃 Did you know? Chlorophyll absorbs strongly in the blue (430 nm) and red (660 nm) regions, but not in green—which is why leaves appear green. Spectrophotometers can measure these specific wavelengths to quantify chlorophyll .

🎯 Choosing the Right Wavelength

For accurate quantification, you should measure at the λmax (wavelength of maximum absorption). This provides:

  • Maximum sensitivity: Small changes in concentration give larger changes in absorbance
  • Minimum interference: Measurements are less affected by small wavelength errors
📊 [Graph: Absorption spectrum showing λmax — to be inserted]

To find λmax, you can perform a wavelength scan (if your instrument allows) or consult published data.

⚠️ Common Pitfalls and How to Avoid Them

Problem Cause Solution
Absorbance > 1.5 Sample too concentrated; Beer-Lambert law may not hold Dilute sample and repeat; ensure A < 1.0 for best accuracy
Negative absorbance Blank has higher absorbance than sample Check blank; might be contaminated or sample less than blank
Poor R² in standard curve Pipetting errors, degraded standards, bubbles in cuvettes Repeat with fresh standards; practice pipetting technique
Bubbles in cuvette Scatter light, giving falsely high absorbance Tap cuvette gently to remove bubbles
Cuvette orientation Fingerprints or scratches on clear sides Always handle cuvettes by the frosted sides; wipe clean with lens paper

Key tip: Always zero the spectrophotometer with a blank between measurements, especially if you change wavelengths .

✏️ Practice Problems

1. Beer-Lambert calculation: A solution of NADH has an absorbance of 0.45 at 340 nm in a 1 cm cuvette. The molar extinction coefficient of NADH at 340 nm is 6220 M⁻¹cm⁻¹. What is the concentration of NADH?

Show solution

A = ε × c × l
0.45 = 6220 × c × 1
c = 0.45 / 6220 = 7.23 × 10⁻⁵ M = 72.3 μM

2. Standard curve: A BSA standard curve gives equation y = 0.85x + 0.02 (y = A₅₉₅, x = mg/mL). An unknown protein sample gives A₅₉₅ = 0.62. What is its concentration? If you diluted your sample 10-fold before assay, what was the original concentration?

Show solution

0.62 = 0.85x + 0.02 → x = (0.62 - 0.02)/0.85 = 0.71 mg/mL (in cuvette)
Original concentration = 0.71 × 10 = 7.1 mg/mL

🇪🇹 Spectrophotometry in Ethiopian Labs

Many Ethiopian universities and research institutions have spectrophotometers, but they may be older models. Tips for success:

  • Know your instrument: Some older models require warm-up time (15-30 minutes) before stable readings
  • Wavelength calibration: Check periodically with known standards (e.g., holmium oxide filter)
  • Cuvette care: Glass cuvettes work for visible light, but quartz is needed for UV (<340 nm)
  • Reagent stability: Some reagents (Bradford, Folin-Ciocalteu) may degrade in heat; store properly

🌱 Measuring Chlorophyll in Teff

Researchers at an Ethiopian university are studying drought tolerance in teff. They need to measure chlorophyll content as an indicator of stress. Using a simple spectrophotometer and the acetone extraction method, they can quantify chlorophyll a and b. This data helps identify drought-tolerant varieties .

📌 Unit Summary

Concept Key points
Beer-Lambert Law A = εcl; absorbance is directly proportional to concentration
Standard curves Plot absorbance vs. concentration for known standards; use equation to find unknowns
Bradford assay 595 nm; Coomassie dye binds protein; fast and sensitive
Phenol-sulfuric acid 490 nm; measures total carbohydrates
Folin-Ciocalteu 765 nm; measures total phenolics (gallic acid equivalents)
Chlorophyll 647 nm and 663 nm; Arnon's equations for chl a and b
Reflection question: You are measuring protein concentration in leaf extracts using the Bradford assay. Your samples give absorbance readings >1.5, and your standard curve is linear only up to 1.0 mg/mL. How would you modify your procedure to obtain accurate results?

📌 Key terms introduced

Spectrophotometry Beer-Lambert law Absorbance (A) Transmittance (T) Molar extinction coefficient (ε) λmax Standard curve Bradford assay Phenol-sulfuric acid Folin-Ciocalteu Arnon's equations Cuvette

✅ Check your understanding

  1. Write the Beer-Lambert law and define each term.
  2. Why is it important to measure absorbance at λmax?
  3. Describe how to construct a standard curve for protein quantification.
  4. What would you do if your sample absorbance is 1.8, but your standard curve is linear only to A = 1.0?
  5. What wavelengths would you measure to quantify chlorophyll a and b?

Discuss your answers in the course forum.

Plant Biochemistry for Horticulture · HORT 202 · Dilla University · Last updated March 2026